Knothole Yoke-moss (Codonoblepharon forsteri)

Key Details

Taxonomic Groups: Non-vascular plant (incl. chromists) > moss > Moss
Red List Status: Endangered (Not Relevant) [EN(nr)]
D5 Status: Included in the baseline Red List Index for England (Wilkins, Wilson & Brown, 2022)
Section 41 Status: (not listed)
Taxa Included Synonym: Zygodon forsteri
UKSI Recommended Name: Codonoblepharon forsteri
UKSI Recommended Authority: (Dicks.) Goffinet
UKSI Recommended Qualifier: (none specified)
Red List Citation: Callaghan, 2023
Notes on taxonomy/listing: in s41 as Zygodon forsteri

Criteria

Question 1: Does species need conservation or recovery in England?
Response: Yes
Justification: At risk of extinction in England due to small number of locations, small population size and continuing decline
Question 2: Does recovery/ conservation depend on species-specific actions?
Response: Yes
Justification: Research is needed to determine effective conservation approaches
Question 3: At a landscape scale, would the species benefit from untargeted habitat management to increase habitat mosaics, structural diversity, or particular successional stages?
Response: No
Justification: The species has a restricted distribution and so is unlikely to benefit from untargeted actions

Species Assessment

Current step on the Species Recovery Curve (SRC): 6. Recovery solutions trialled
Recovery potential/expectation: Unknown
National Monitoring Resource: Opportunistic - insufficient
Species Comments: GB RL criteria: B1ab(iii,v), B2ab(iii,v), C2a(i). Southern-temperate Suboceanic species.

Key Actions

Key Action 1

Proposed Action: Develop and implement national monitoring plan to ensure comprehensive and up-to-date information is available on range size, population size, occupied sites and threatening factors in England

Action targets: 3. National Monitoring Plan agreed and implemented

Action type: Targeted monitoring

Duration: 1 year

Scale of Implementation: ≤ 5 sites

High priority sites: Burnham Beeches SSSI, Epping Forest SSSI, The New Forest SSSI

Comments: A species with low recovery potential where the main aim is to maintain/protect populations

Key Action 2

Proposed Action: Assess feasibility and develop a plan for reintroduction to South Somerset and Worcestershire

Action targets: 6. Recovery solutions trialled

Action type: (Re-)introduction

Duration: 1 year

Scale of Implementation: ≤ 5 sites

High priority sites:

Comments: Historically known from South Somerset and Worcestershire, where reintroduction may be a possibility, using direct translocation of spores from a donor site.

Key Action 3

Proposed Action: Undertake tree veteranisation trials to develop methods for creating suitable niche space for the species

Action targets: 6. Recovery solutions trialled

Action type: Scientific research

Duration: 3-5 years

Scale of Implementation: ≤ 5 sites

High priority sites: Epping Forest SSSI, Burnham Beeches SSSI, The New Forest SSSI

Comments:

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Acknowledgment:
Data used on this website are adapted from Threatened species recovery actions 2025 baseline (JP065): Technical report and spreadsheet user guide (Natural England, 2025). Available here.